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Fig. 3. Intergeneric hybrid (2n = 5x = 35; ABDPP genomes) between bread wheat (2n = 6x = 42; AABBDD) wheatgrass, Agropyron cristatum (2n = 4x = 28; PPPP), and genotype-induced homoeologous chromosome pairing. (A) Bread wheat var. Fukuhokomugi (left), crested wheatgrass (right), and their hybrid (center). Note the intermediate phenotype of the hybrid. (B) Meiotic metaphase in the hybrid showing 1 III + 10 II + 12 I. The genotype of the grass parent induces higher pairing than is expected based on chromosome homology.