Published in Crop Sci. 44:1615-1621 (2004).
© 2004 Crop Science Society of America
677 S. Segoe Rd., Madison, WI 53711 USA
CROP PHYSIOLOGY & METABOLISM
Nitrogen Rate and Mowing Height Effects on Turf-Type Buffalograss
K. W. Franka,*,
R. E. Gaussoinb,
T. P. Riordanb,
R. C. Shearmanb,
J. D. Fryc,
E. D. Miltnerd and
P. G. Johnsone
a Dep. of Crop and Soil Sci., Michigan St. Univ., East Lansing, MI 48824
b Dep. of Agronomy and Horticulture, Univ. of Nebraska, Lincoln, NE 68583
c Div. of Horticulture, Kansas State Univ., Manhattan, KS 66506
d Washington St. Univ., Puyallup Research Stn., 7612 Pioneer Way East, Puyallup, WA 98373
e Dep. of Plants, Soils, and Biometeorology, Utah St. Univ., 4820 Old Main Hill, Logan, UT 84322
* Corresponding author (frankk{at}msu.edu).
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ABSTRACT
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Buffalograss [Buchloë dactyloides (Nutt.) Engelm.] is a turfgrass species with reduced irrigation and fertilization requirements, relative to traditional turfgrasses. Interest in reducing the amount of chemicals and water applied to turfgrass has resulted in increased efforts to improve buffalograss as a turfgrass. As interest in using the new turf-type buffalograss cultivars has increased, the need for nitrogen rate and mowing height recommendations supported by research have become necessary. Cody, Texoka, 378, and NE 91-118 buffalograss cultivars were planted at sites located in Nebraska, Kansas, and Utah to determine nitrogen rate and mowing height effects on buffalograss quality, color, density, and clipping yield. There were significant cultivar x nitrogen rate interactions at all sites for quality, color, and density. Regardless of cultivar, nitrogen rate x year interactions at all sites revealed that the 98 kg N ha1 rate sustained quality, color, and density over the 3-yr evaluation period, while lower nitrogen rates had decreased quality, color, and density. The 195 kg N ha1 rate improved quality over the 3-yr period but also had the highest clipping yields. Regardless of the cultivar selected, the data support the general mowing height recommendation for turf-type buffalograss of 5.0 to 7.5 cm. For golf course fairways maintained at a mowing height of 2.5 cm, the vegetatively established cultivars (NE 91-118 and 378) are the best choice to achieve high turfgrass quality.
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INTRODUCTION
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BUFFALOGRASS is a warm season, C4 grass native to the Great Plains of North America (Wenger, 1943). Exceptional drought tolerance, good heat resistance, and cold tolerance enable buffalograss to be distributed across the Great Plains (Beard, 1973; Beetle, 1950; Savage, 1933; Wenger, 1943).
Interest in buffalograss as a turfgrass species is primarily due to its water conservation and minimal maintenance characteristics (Pozarnsky, 1983; Riordan, 1991; Wu and Harivandi, 1989). Recently developed turf-type buffalograss cultivars have been used on home lawns, golf courses, public grounds, and highways (Cline, 1994; Falkenberg-Borland and Butler, 1982; Fry, 1995; Wu and Harivandi, 1989).
Buffalograss has been described in numerous Extension and trade publications as a low maintenance turfgrass that has reduced irrigation, nitrogen, and mowing requirements and few disease and insect pests (Pozarnsky, 1983; Leuthold et al., 1991; Riordan, 1991; Wu et al., 1991), but scientific-based information supporting these claims is lacking. Management recommendations appear to be based on observation and not replicated research trials. Extension bulletins suggest the desired use and quality of buffalograss determines the choice of mowing height and frequency (deShazer et al., 1991; Leuthold et al., 1991). Mowing height recommendations for buffalograss range between 5 and 10 cm (deShazer et al., 1992).
Research on buffalograss in Colorado indicated that moderate levels of nitrogen fertilization improve buffalograss growth, quality, color, and density (Falkenberg, 1982). However, the buffalograss cultivar used in Falkenberg's research was not specified and the results achieved may certainly differ from the new, improved turf-type buffalograss cultivars. Current fertilizer recommendations range from 0 to 98 kg N ha1 yr1 (deShazer et al., 1991; Leuthold et al., 1991; Riordan, 1991). Excessive or frequent fertilizer applications are reported to encourage weed interference and have little, if any, positive effect on buffalograss quality (deShazer et al., 1991; Harivandi and Wu, 1995; Leuthold et al., 1991; Riordan, 1991).
Although general buffalograss management recommendations pertaining to fertilization and mowing are available, research on turf-type buffalograss management is limited. With this in mind, this research was conducted to determine nitrogen rate and mowing height effects on buffalograss turfgrass quality, color, density, and clipping yield.
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MATERIALS AND METHODS
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Two vegetative cultivars, 378 and NE 91-118, and two seeded cultivars, Cody and Texoka, were planted in 1995 at three sites: the John Seaton Anderson Turfgrass Research Facility located at the University of Nebraska Agricultural Research and Development Center near Mead, NE; Kansas State University Rocky Ford Turfgrass Research Center at Manhattan, KS; and the Utah State University Greenville Research Farm at Logan, UT. The soil types at the Nebraska, Kansas, and Utah sites were a Tomek silty clay loam (fine montmorillonitic, mesic Typic Argiudoll), a Chase silt loam (fine, montmorillonitic, messic, Aquic, Arquicolls), and a Millville silt loam (coarse-silty, carbonatic, mesic Typic Haploxepolls), respectively. Soil samples were taken before establishment at all sites and tested for pH, P, and K. At the Nebraska site: pH = 7.0, P = 71 µg g1 (Bray P1 extractable P), and K = 413 µg g1. At the Kansas site: pH = 6.8, P = 50 µg g1 (Bray P1 extractable P), and K = 471 µg g1. At the Utah site: pH = 8.0, P = 30 µg g1 (Olsen extractable P), and K = 471 µg g1.
The same planting procedures were used at the three locations. Vegetative plugs of NE 91-118 and 378, 5 cm in diam., were planted on 30-cm centers and Cody and Texoka were seeded at 98 kg burs ha1. Cody and Texoka seed were incorporated into the soil to a depth of approximately 1 cm using a leaf rake. After planting, a cultipacker was rolled over the plots to facilitate plug and seed to soil contact. Starter fertilizer (19N-11P-4K) was applied at 49 kg N ha1 and simazine herbicide (6-chloro-N, N'-diethyl-1,3,5-triazine-2,4-diamine) was applied at 1.12 kg a.i. ha1 at planting. Four weeks after planting (WAP), sulfur coated urea fertilizer (25N-0P-8K) was applied at 49 kg N ha1 and at 8 WAP urea (46N-0P-0K) was applied at 24 kg N ha1. Plots were mowed periodically at 6.4 cm and irrigation was applied as necessary to maintain a moist seedbed during the establishment year.
Buffalograss was successfully established at the Nebraska and Kansas sites in 1995. At the Utah site, 378 and NE 91-118 failed to cover the plots in 1995, but the seeded cultivars were successfully established. The transport of plugs to Utah, which included washing of soil from the plugs, was believed to be a factor slowing establishment. Because of negligible survival of 378 at the Utah site, the decision was made to abandon 378 and concentrate on establishment of NE 91-118. Additional NE 91-118 plant material was transported to the Utah site and planted in May of 1996. By the end of the 1996 growing season, the NE 91-118 plots had at least 70% buffalograss cover.
Preemergence herbicides were applied each year from 1996 to 1998. At the Nebraska and Kansas sites, pendimethalin [N-(1-ethylpropyl)-3,4-dimethyl-2,6-dintrobenzenamine] was applied at 3 kg ai ha1 in 1996 and dithiopyr [S,S-dimethyl-2-(difluoromethyl)-4-(2-methylpropyl)-6-(trifluoromethyl)-3,5-pyridinedicarbothioate] was applied at 0.56 kg ai ha1 in 1997 and 1998. At the Utah site simazine was applied at 1.12 kg ai ha1 in May 1996 and pendimethalin was applied at 3 kg ai ha1 in 1997 and 1998.
In 1996, mowing heights and nitrogen treatments were initiated. Mowing heights were 2.5, 5.0, and 7.5 cm. Plots were mowed weekly with clippings removed. Nitrogen was applied in two equal applications with the first application in early June and the second application in mid-July, 6 wk after the first application. A polymer coated fertilizer (36N-1P-6K) was used to apply total nitrogen amounts of 24, 49, 98, and 195 kg N ha1 yr1. In addition, an untreated control was included. Immediately after fertilizer applications, plots were irrigated with approximately 1.3 cm water. After adjusting for precipitation at each site, 5.0 cm of water was applied every 2 wk throughout the research.
Turfgrass quality, color, and density were rated visually on a scale of 1 to 9 with 1 = poor, 6 = acceptable, and 9 = excellent. The rating scale for density was 1 = least and 9 = most. The rating scale for color was 1 = straw brown, 6 = acceptable, and 9 = dark green. Ratings were taken every 2 wk after the first nitrogen application and continued until 6 wk after the second nitrogen application. Clippings were harvested 4 wk after each fertilizer application, dried at 60°C for 48 h and dry weights recorded.
Experimental design for each site was a randomized incomplete block design and treatment design was a split-split-plot. Main plots were buffalograss cultivars, split plots were mowing heights, and split-split plots were nitrogen rates. Statistical analysis indicated site effects; therefore, data were analyzed separately by site. Treatment differences were tested using Proc Mixed statistical analysis (SAS Institute Inc., 1997). Means were separated using Fisher's Protected LSD procedure.
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RESULTS AND DISCUSSION
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The results of analysis of variance for turfgrass quality are presented in Tables 1, 2, and 3 for the Nebraska, Kansas, and Utah sites, respectively.
Nitrogen Rate Response
Cultivar x Nitrogen Rate
At the Nebraska site, there were cultivar x nitrogen rate interactions for turfgrass quality in each year from 1996 to 1998 (Table 1). At 4 wk after the second nitrogen application in 1996, 378 had the highest quality when no nitrogen was applied (Table 4). Four weeks after the second nitrogen application in 1997, Cody, 378, and NE 91-118 had the highest quality when no nitrogen was applied (Table 4) and at 4 wk after the second nitrogen application in 1998, 378 and Cody had the highest quality when no nitrogen was applied (Table 4). Only in 1996 were any of the quality ratings greater than the acceptable rating of 6. For 378, the quality rating from 1996 to 1998 declined from 6.1 to 4.0. At the Kansas and Utah sites, turfgrass quality for the untreated control never reached the acceptable rating of 6 (Table 5).
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Table 4. Mean buffalograss quality for the cultivar x nitrogen rate interaction at 4 wk after the second nitrogen application in 1996, 1997, and 1998 at the Nebraska site.
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Table 5. Mean buffalograss quality for the cultivar x nitrogen rate interaction at 4 wk after the second nitrogen application in 1996 at the Utah site and 1997 at the Kansas site.
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In Nebraska, 4 wk after the second nitrogen application in 1997 and 1998, 378, NE 91-118, and Cody had acceptable turfgrass quality ratings at the 98 kg N ha1 rate (Table 4). In 1997 at the Kansas site, NE 91-118 required 49 kg N ha1 for acceptable quality, all other cultivars required 98 kg N ha1 (Table 5). At the Utah site in 1996, NE 91-118 and Cody required 98 kg N ha1 for acceptable quality (Table 5). At the Nebraska and Utah sites Texoka required 195 kg N ha1 to reach acceptable turfgrass quality.
The cultivar x nitrogen rate interaction was not always significant at all sites throughout 1996 to 1998; therefore, direct comparisons among sites are limited. Environmental conditions and soil characteristics at the sites may favor one cultivar over another. Identification of the factors that are the primary determinants of turfgrass quality at the sites is beyond the scope of this research. However, it should be recognized that the cultivar response to nitrogen rates did differ among sites and site differences could potentially influence management recommendations.
Although the response of specific cultivars to nitrogen rates at the sites varied, the overall turfgrass quality responses to nitrogen were similar. As nitrogen rate increased, turfgrass quality improved and acceptable quality ratings for all cultivars, except Texoka, required 98 kg N ha1. The primary difference among the cultivars at the sites was at the low nitrogen rates where quality was often unacceptable. Texoka required 195 kg N ha1 to reach acceptable turfgrass quality ratings. There are several explanations as to why Texoka required a higher nitrogen rate to achieve acceptable turfgrass quality ratings. Although used as a minimal maintenance turfgrass, Texoka was developed for use as a forage grass on rangelands (Voigt et al., 1974). In comparison to the improved turf-type seeded cultivars, such as Cody, Texoka produces a turf stand that is less dense. The density ratings for Texoka, especially at the 2.5-cm mowing height, were often very low throughout the research (data not presented) and the lack of density definitely influenced the quality ratings.
Nitrogen Rate x Year
There was a nitrogen rate x year interaction for turfgrass quality, color, and density at all sites. The quality response to nitrogen rates from 1996 to 1998 was similar for all sites. Data will be presented for the Nebraska site only. Quality declined from 1996 to 1998 for the 0, 24, and 49 kg N ha1 rates (Table 6). There were no differences in quality from 1996 to 1998 for the 98 kg N ha1 rate. Quality improved from 1996 to 1998 at the 195 kg N ha1 rate. The 98 kg N ha1 rate sustained turfgrass quality over three years, while lower nitrogen rates did not.
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Table 6. Mean buffalograss quality and color for the nitrogen rate x year interaction at 4 wk after the second nitrogen application at the Nebraska site.
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For turfgrass color, the nitrogen rate x year interaction revealed a similar trend as color ratings declined over time at the 0, 24, and 49 kg N ha1 rates. There was variability among the sites as to the year when color declined or increased for the different nitrogen rates, but overall the trends were similar at all sites. For the 0, 24, and 49 kg N ha1 rates, color declined from 1996 to 1998 at all sites (Data presented from Nebraska site only, Table 6). At the 98 kg N ha1 rate there were no differences in color among years. At the Kansas site there were no differences in color at the 98 kg N ha1 rate in 1996 and 1997, but color ratings declined from 6.3 in 1997 to 5.8 in 1998. At the Utah site, color was highest in 1997 at the 98 kg N ha1 rate and there were no differences in 1996 and 1998. The 98 kg N ha1 rate sustained color over the three year period and the 195 kg N ha1 rate improved color. The 0, 24, and 49 kg N ha1 rates resulted in decreased color over the 3-yr period at the Utah site.
Turfgrass density responses over the 3-yr period closely resembled the quality and color responses. There was a nitrogen rate x year interaction for turfgrass density at all sites (data not presented). The interaction revealed the 98 and 195 kg N ha1 rate sustained density over the 3-yr period but the 0, 24, and 49 kg N ha1 rates had declining density over time.
Clipping Yield
The 195 kg N ha1 rate had the highest turfgrass quality but for some cultivars also produced greater amounts of clippings than the 98 kg N ha1 rate. The nitrogen rate x cultivar interaction for clipping weights was significant at the Nebraska and Kansas sites in 1997 and 1998. At the Utah site, N rate was significant 1996 to 1998 with the 195 kg N ha1 rate having the highest clipping weight (data not presented). At the Nebraska site in 1997, there were no differences in clipping weights between the 98 and 195 kg N ha1 rate for NE 91-118 and 378, but clipping weights were highest at the 195 kg N ha1 rate for Cody and Texoka (Fig. 1)
. At the Kansas site, the results were similar although the clipping weight response to nitrogen rates for vegetative (NE 91-118 and 378) and seeded (Cody and Texoka) selections was not as distinct (data not presented). There was no difference in clipping weights between 98 and 195 kg N ha1 for NE 91-118, but for 378, Cody, and Texoka the 195 kg N ha1 rate had the highest clipping weights. Although the 195 kg N ha1 rate had the highest quality it also produced the highest amount of turfgrass clippings. If the desire is to manage buffalograss as a minimal maintenance turfgrass with infrequent mowing, the 195 kg N ha1 rate would not be favorable because of excessive clipping production.

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Fig. 1. Clipping weights for the cultivar x nitrogen rate interaction at 4 wk after the second nitrogen application in 1997 at the Nebraska site. Significance at 0.05 level of probability among cultivars at the same nitrogen rate is indicated with an * or ns (not significant) at each nitrogen rate.
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Nitrogen Rate Results
The results of the cultivar x nitrogen rate and nitrogen rate x year interactions revealed that the 98 kg N ha1 rate provided acceptable turfgrass quality for all cultivars, except Texoka, and sustained the quality over 3 yr. Although the 195 kg N ha1 rate steadily improved quality over the 3-yr period and resulted in the highest turfgrass quality among the nitrogen rates, it also resulted in the highest clipping yields. The higher clipping yields produced by the 195 kg N ha1 rate and the relatively high amount of nitrogen applied, render this rate undesirable when considering the desire for reduced inputs in a buffalograss management program. Leuthold et al. (1991) recognized this fact when recommending that nitrogen levels for buffalograss not exceed 98 kg N ha1 yr1. Although this research contradicts the perception that buffalograss does not respond favorably to nitrogen levels in excess of 98 kg N ha1 yr1, it does support the recommendation of 98 kg N ha1 yr1 for reduced input turf-type buffalograss maintenance.
Mowing Height Response
Differences among Cultivars
The cultivar x mowing height interaction for turfgrass quality was significant for each year from 1996 to 1998 at the Nebraska site, 1997 and 1998 at the Kansas site, and 1996 and 1997 at the Utah site (Tables 1, 2, and 3). The overall results from each site were similar. In 1997 at the Nebraska and Kansas sites at 4 wk after the second nitrogen application, NE 91-118, 378, and Cody had the highest quality at the 2.5-cm mowing height (Table 7). At the Utah site, NE 91-118 had the highest quality at the 2.5-cm mowing height; however, 378 was not tested at this location (Table 7). At the 2.5-cm mowing height at all locations, Texoka consistently had the lowest quality ratings; but, at the Nebraska site in 1998 and at the Kansas site in 1997 and 1998 Texoka's quality was not different than Cody (Tables 7 and 8).
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Table 7. Mean buffalograss quality for the cultivar x mowing height interaction at 4 wk after the second nitrogen application in 1997 at the Nebraska, Kansas, and Utah sites.
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Table 8. Mean buffalograss quality for the cultivar x mowing height interaction at 4 wk after the second nitrogen application in 1998 at the Nebraska site and Kansas sites.
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At the Nebraska site there were no differences in quality among the cultivars at the 5.0-cm mowing height in 1997. At the Nebraska site in 1996 and 1998, NE 91-118, 378, and Cody had the highest quality at the 5.0-cm mowing height; however, only 378 had a higher quality rating than Texoka (Table 8, data not presented from 1996). At the Kansas and Utah sites in 1997 and 1998, NE 91-118, 378, and Cody had the highest quality at the 5.0-cm mowing height but only NE 91-118 had a higher quality rating than Texoka (Tables 7 and 8). Overall, these results indicate that there were very few differences in turfgrass quality among the cultivars at the 5.0-cm mowing height. The result that was similar among years and sites was that either 378 or NE 91-118 had higher quality ratings than Texoka, and Cody and Texoka were equal in turfgrass quality.
For the 7.5-cm mowing height at the Nebraska site there were no differences among cultivars in 1996 and 1997 (Table 7). In 1998, Cody, Texoka, and 378 had the highest numerical quality values, however the only statistical difference in quality was between 378 and NE 91-118 (Table 8). At the Kansas and Utah sites there were no differences in quality among cultivars at the 7.5-cm mowing height in 1997 and 1998 or 1996 and 1997, respectively (Tables 7 and 8). In almost all cases at all sites there were no differences in quality among the cultivars at the 7.5-cm mowing height. Unlike the 2.5- and 5.0-cm mowing heights, Texoka performed equally to other cultivars at the 7.5-cm mowing height.
Mowing Height Results for Cultivars
Although mowing height results for each cultivar are not entirely consistent for all locations and years, the following conclusions can be drawn.
NE 91-118 appears to be adapted to a wide range of mowing heights while still achieving acceptable turfgrass quality. The result that was consistent was that among mowing heights NE 91-118 had lower quality ratings at the 7.5-cm mowing height. This was true for all sites in 1997 and for the Kansas site in 1998 (Tables 7 and 8). Turfgrass quality at the 7.5-cm mowing height was lower than the other mowing heights because of stolons lying on top of the canopy and generally detracting from the overall uniformity of the turfgrass stand. This response would certainly not make the 7.5-cm mowing height unacceptable for low maintenance sites but for high profile areas this appearance may be undesirable. The 2.5-cm mowing height would only be recommended for golf course fairways or other intensively maintained sites.
For 378 and Cody, the 5.0- and 7.5-cm mowing height resulted in the highest quality at the Nebraska site in 1998 and the Utah site in 1997. At the Kansas site there were no differences in quality among mowing heights for 378 and Cody, nor were there any differences at the Nebraska site in 1997. For Texoka, the 5.0- and 7.5-cm mowing heights were clearly preferred to obtain the highest quality.
Our data supports the general mowing height recommendation for turf-type buffalograss of 5.0 to 7.5 cm. The data also support tolerance to lower mowing heights (i.e., 2.5 cm) by cultivars such as NE 91-118 and 378. These results substantiate past views expressed in extension publications on the recommended mowing height for buffalograss (deShazer et al., 1991; Leuthold et al., 1991).
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CONCLUSIONS
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Because of differences in cultivar response to the nitrogen and mowing height treatments at the three sites, making specific management recommendations for the buffalograss cultivars is difficult. However, several nitrogen rate and mowing height effects were consistent among sites. The nitrogen rate x year interaction was significant at all sites for buffalograss quality, color, and density. The 98 kg N ha1 yr1 rate sustained quality, color, and density over the 3-yr period, but lower nitrogen rates resulted in decreased quality, color, and density. The 195 kg N ha1 yr1 rate increased quality, color, and density over the 3-yr period but also had the highest clipping weights, especially for the seeded cultivars Cody and Texoka. Based on our results, the 98 kg N ha1 yr1 rate is sufficient to provide acceptable turfgrass quality and color for the cultivars tested. The higher nitrogen rate (195 kg N ha1 yr1) enhanced color and overall quality, but this rate would seem to contradict the use of buffalograss as a low maintenance turfgrass with relatively low nitrogen and mowing frequency requirements. If a high level of management is desired, including frequent mowing due to high clipping production, the 195 kg N ha1 yr1 rate would be an acceptable choice. For situations where a low input management program with modest nitrogen rates and infrequent mowing are desired, the 98 kg N ha1 rate is the best choice.
Mowing heights in the range of 5.0-7.5 cm provide acceptable turf quality for the cultivars tested. The use of lower (98 kg ha1 yr1) nitrogen rates and higher mowing heights (5.07.5 cm) provide a low maintenance buffalograss turf of acceptable quality. Although NE 91-118 and 378 responded favorably to both the higher (195 kg ha1 yr1) nitrogen rate and low (2.5 cm) mowing height, these levels of maintenance are only recommended when these cultivars are utilized for golf course fairways or other high maintenance turf.
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ACKNOWLEDGMENTS
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The authors express thanks to the United States Golf Association for providing financial support for this research. The authors would also like to thank Leonard Wit, Jeff Witkowski, and Alan Zuk for their assistance in plot maintenance.
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NOTES
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Contribution of Kansas Agric. Exp. Stn. No. 04-049-J.
Received for publication July 14, 2003.
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REFERENCES
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- Falkenberg, D.F. 1982. Buffalograss, blue grama, and fairway wheatgrass for dryland turf. M.S. Thesis. Colorado State Univ., Fort Collins.
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- Riordan, T.P. 1991. Buffalograss. p. 1214. Grounds Maintenance. February.
- SAS Institute Inc. 1997. SAS/STAT Software: Changes and enhancements through release 6.12. SAS Inst., Cary, NC.
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- Voigt, P.W., W.R. Kneebone, J.R. Harlan, and R.M. Ahring. 1975. Registration of Texoka buffalograss. Crop Sci. 15:885.[Free Full Text]
- Wenger, L.E. 1943. Buffalo Grass. Kans. Agric. Exp. Stn. Bull. 321.
- Wu, L., and A. Harivandi. 1989. Buffalograss: Promising, drought-resistant- and here now. Golf Course Manage. 57(4):4245.
- Wu, L., D. Huff, and A. Harivandi. 1991. Buffalograss as a low-maintenance turf. p. 2325. California Agriculture. March-April.
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