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a Faculty of Agriculture, Utsunomiya Univ., 350 Mine-Machi, Utsunomiya, 321-8505 Japan
b National Agriculture Research Center for Hokkaido Region, Hitsujigaoka 1, Sapporo, 062-8555 Japan
c National Institute of Crop Science, Kannondai 2-1-18, Tsukuba, Ibaraki, 305-8518 Japan
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