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An aberrant plant with an extreme cup leaf expression was found in a population of true breeding cup leaf plant~;. The mutant was called super cup. Cytological analysis revealed that the super cup mutant was a trisomic plant. A crossing program with known translocations identified the replicated chromosome as chromosome 4 of the A genome. Genetic tests determined that the cup leaf locus is independent of the replicated chromosome; and the super cup phenotype is due to an interaction of the cup leaf locus with the replicated chromosome.
2 Research Geneticist, Crops Research Division, ARS, USDA.
Received for publication August 27, 1965.
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